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M Killifish
The "M" Killifish
Species List
Nimbapanchax leucopterygius
Lola -
The old "Roloffia Aphyosemion / Archaphyosemion maeseni turned out to have been described from a type specimen that was really an Epiplatys. This is the new name for that fish.
Rivulus (Laimosemion) mahdiaensis
desc biotope
karyo -  (2n=38) Guy_97-5

Rivulus mahdiaensis

Discovered in 1995 by W. Suijker and Y. Suijker-Jansen who also found another location 1997 with F. and M. Vermeulen; no other locations have been found since. Described by W. Suijker and G. Collier in 2006. Unusually low number of chromosomes with a rather large pair of arcocentric elements (shown with arrows). Uneven sex ratios are common, not difficult otherwise. From cool acidic soft "black" water similar to African killifish blackwater biotopes for example, Diopteronspecies.
Funduls majalis

Funduls majalis

Nothobranchius makondorum
Melela_River -
Molocue_River -

Nothobranchius makondorum

Aphyosemion maculatum
ABDEK_12-495 -   Bisun -   DNA_1_LOLO-1 -
GEMLH_16-28 -   PEG_94 -  aff -

Aphyosemion maculatum

Fundulopanchax marmoratus
Bakossi -   Ituka -
Mbonge -   Mundemba -

Fundulopanchax (Pauciradius) marmoratus

Rivulus (Laimosemion) mabura

Rivulus mabura

Hypsolebias macaubensis

Hypsolebias macaubensis

Rachovia maculipinnis

Rachovia maculipinnis

Anablepsoides micropus

Anablepsoides micropus

Cyprinodon macularius

Cyprinodon macularius

Pantanodon madagascariensis

Pantanodon madagascariensis

Platypanchax modestus
Ntoroko

Platypanchax modestus

Aphanius maeandricus

Aphanius maeandricus

Hypsolebias mediopapillatus

Hypsolebias mediopapillatus

Rivulus mejiai

Rivulus mejiai

Cyprinodon meeki

Cyprinodon meeki

Paraphanius mento
Zengen Domasa
Ein_Feshka Elbistan

Paraphanius mento

Nothobranchius melanospilus
Mbezi -  Mbezi river floodplains Mwakanga Ramisi_KE_01-12
TSTS_2010-16 Vigwaza aff -  N. affinis melanospilus

Nothobranchius melanospilus

Epiplatys mesogramma
cf -  Z86/15
Bangui

Epiplatys mesogramma

Lacustricola margaritatus

Lacustricola margaritatus

Lacustricola margaritatus, a new species inhabiting small streams and swamps in the Lake Victoria basin in north-western Tanzania and southern Uganda, and the Lake Kyoga basin in central Uganda, is described. Lacustricola margaritatus is a small species with a moderately deep body, moderate dimorphism and pronounced dichromatism. It is distinguished from all other Procatopodidae by the following unique combination of characters: live male body colour pattern with vertically-elongated iridescent light blue patches at scale centres, forming a striped appearance of dotted longitudinal lines on the flanks, particularly evident in the two or three series below the mid-longitudinal line; male having deeply coloured unpaired fins with orange-brown in the proximal and median parts and a narrow black distal band; male with a yellow base along the pectoral fin; female with dark grey scale margins and dark grey patches on scales along mid-longitudinal series creating a narrow dark grey stripe; both sexes showing inconspicuous postopercular blotch; and in both sexes, the cephalic sensory system is entirely situated in open grooves at all levels. The new species has previously often been misidentified as L. pumilus, originally described as inhabiting the Lake Tanganyika basin in north-eastern Zambia, or 'L.' centralis, from the Lake Rukwa basin in south-western Tanzania. Lacustricola margaritatus differs from the above two species by morphometric and meristic characters, body and fin colouration, and in arrangement of the cephalic sensory system.
Cynodonichthys magdalenae
Ibague -
Santandercito -  Tocaima -

Cynodonichthys magdalenae

The highland form has an orange band in the tail, the lowland form has a white bar in the tail.
Hypsolebias magnificus
Itacarambi -

Hypsolebias magnificus

Hypsolebias marginatus
Barro_Alto -

Hypsolebias marginatus

Simpsonichthys margaritatus

Simpsonichthys margaritatus

Nothobranchius (N.mic) microlepis
Tana_River_South -

Nothobranchius (N.mic) microlepis

Aphyosemion mimbon
G02 -   Akoga -  BBS_99-22 -   Bicum_2018-15 -   GCC_13 -
GEB -   GEGVPO_2018-17 -   GEMG_11-24 -   GEMHS -   GEMLB_02-3 -   GEMLC_4-7 -   GHG_83-1 -    GWW_86-30 -    LEC -   NKinnen_DNA_01 -

Aphyosemion mimbon

Leptolebias minimus
Inoa -
Seropedica -

Notholebias minimus

Formerly:
Cynopeocelius ladigesi
Cynloebias minimus
Leptolebias minimus
Aphyosemion microphtalmum

Aphyosemion microphtalmum

Fundulopanchax mirabilis
Mbio -
Nchemba -

Fundulopanchax mirabilis

Anatolichthys meridionalis

Anatolichthys meridionalis

Profundulus  mixtlanensis

Profundulus mixtlanensis

Nothobranchius mkuziensis

Nothobranchius mkuziensis

Aphyosemion malumbresi
GEMLCG_2007-50

Aphyosemion malumbresi

Aphyosemion melanogaster
Lobe -
kribi -

Aphyosemion melanogaster

Cynodonichthys = MGD milesii

Cynodonichthys milesii = MGD

Cyprinodon salinus milleri

Cyprinodon salinus milleri

Cynopoecilus melanotaenia

Cynopoecilus melanotaenia

 Aphyosemion  melinoeides
Akok -
Fenda -

Aphyosemion melinoeides

Austrolebias melanoorus

Austrolebias melanoorus

Lacustricola maculatus

Lacustricola maculatus

Plataplochilus miltotaenia
Gabon
Lambarene

Plataplochilus miltotaenia

Nothobranchius milvertzi
Lushiba_Chienge -

Nothobranchius milvertzi

Nothobranchius moameensis

Nothobranchius moameensis

Leptolebias marmoratus

Leptolebias marmoratus

Plataplochilus mimus

Plataplochilus mimus

Aphyosemion mandoroense

Aphyosemion mandoroense

Aphyosemion montealenense

Aphyosemion montealenense

Aphyosemion mengilai

Aphyosemion mengilai

Cynodonichthys monikae

Cynodonichthys monikae

Fundulopanchax moensis
Bakebe -   Bator -
Eyang -   Kendem -   Manyemen -

Fundulopanchax moensis

Cyprinodon nevadensis mionectes

Cyprinodon nevadensis mionectes

Hylopanchax moke

Hylopanchax moke

Epiplatys dageti monroviae
Harbel -
Robertsfield -   photos_by -

Epiplatys dageti monroviae

Callopanchax monroviae
CI2012 -  Harbel -
Paynesville_97 -   Robertsport -    red -

Callopanchax monroviae

Austrolebias monstrosus

Austrolebias monstrosus

Cynopoecilus multipapaillatus

Cynopoecilus multipapaillatus

Aphyosemion marginatum
Bengui -
Bifoun -   G80-13 -

Aphyosemion marginatum

Rivulus (Atlantirivulus) maricensis

Rivulus maricensis

Atlantirivulus maricensis: UFRJ 9822, holotype, male, 33.4 mm SL: Brazil: Rio de Janeiro: Maricá.
Rivulus (Kryptolebias) marmoratus
Aruba -
Dangriga -

Rivulus marmoratus

(formerly 'Rivulus marmoratus')
Empetrichthys merriami

Empetrichthys merriami

Lacustricola moeruensis

Lacustricola moeruensis

Hypsolebias martinsi

Hypsolebias martinsi

Hylopanchax multisquamatus

Hylopanchax multisquamatus

Nothobranchius malaissei "Kabiasha CD 16/18"Kabiasha CD 16/18
Kabiusha -
Sange_DRCH_2008-06 -

Nothobranchius malaissei

Nothobranchius matanduensis

Nothobranchius matanduensis

Paraphanius mentoides

Paraphanius mentoides

Aphyosemion mitemelense
Esong_Rio_Misee -  GEAV_14-7 -
GEGVPO_2018-15 -   GEMLC_04-05 -

Aphyosemion mitemelense

"Aphyosemion mitemelense can be distinguished from both described species of the ‘A.’ herzogi species group by the male colour pattern of body and unpaired fins. In A. mitemelense the caudal fin has several more or less horizontal relatively wide dark red bands on centre and lower part of fin vs. whole fin with many red interradial lines on blue centre and lower part of fin in ‘A.’ bochtleri and two red lines edging a broad yellow band on fin centre in ‘A.’ herzogi. Background of anal fin green to blue green in A. mitemelense vs. yellow to orange, at least basally, in the other two species.

Aphyosemion mitemelense can be distinguished from neighboring undescribed species from the ‘A.’ herzogi group by having a greenish appearance and a green anal fin with irregular red blotches and/or stripes and a large dark red spot at the base of pectoral fin vs. a reddish appearance and an orange anal fin with regular interradial lines and lacking a red spot at the base of pectoral fin in populations on the right bank of the Mitemele River in southern Equatorial Guinea; by lacking a yellow horizontal band at the base of anal fin vs. having such a band in a population that occur sympathetic near the village of Mveayong in the extreme southeast of Equatorial Guinea. Aphyosemion mitemelense differs from populations between Ngolensok and Acurenam in southeastern Equatorial Guinea by absence of more or less regular red interradial lines in caudal fin centre, lower half of caudal greenish versus yellow and upper part of fin with many irregular red spots versus most part of upper half of caudal fin with red interradial lines; it differs also by the presence of broad red blotches, stripes or bands on greenish or blue-greenish anal fin versus more narrow, short red interradial lines on yellow or orange anal fin."
P17-18 "Three New Killifish Species (Cyprinodinriformes: Nothobranchiidae) From Equatorial Guinea" (PDF), Francisco J. Malumbres, Rainer Sonnenberg & Jouke R. Van der Zee, Oct 2022
Micropanchax macrophthalmus

Micropanchax macrophthalmus

Rivulus (Cynodonichthys) montium
Nombre_de_Dios -

Rivulus montium

was, Rivulus.
MTN

Oryzias matanensis

Anatolichthys marassantensis

Anatolichthys marassantensis

Lacustricola matthesi

Lacustricola matthesi

Epiplatys multifasciatus

Epiplatys multifasciatus

Austrolebias minuano

Austrolebias minuano

Aphyosemion musafirii

Aphyosemion musafirii

Lacustricola myaposae
Mkuze_ZA_19-1 -
Lake_Sibaya -

Lacustricola myaposae

Nimbapanchax melanopterygius
Cavally_River -

Nimbapanchax melanopterygius

Austrofundulus myersi
aff -

Austrofundulus myersi

Simsonichthys myersi

Xenurolebias myersi

Poropanchax myersi
aff -
ci -  maps -

Poropanchax myersi

Fundulopanchax gardneri mamfense

Fundulopanchax gardneri mamfense


Sorted by Code
MAA   Cyprinodon maya
MAB   Austrolebias (=AFI) mabelis
MAC   Epiplatys (=SIN) macrostigma
MAE   Archiaphyosemion (Nimbapanchax) maeseni
MAG   Aphyosemion (=CHR) margaretae
MAH   Rivulus (Laimosemion) mahdiaensis
MAI   Aplocheilus panchax mattei
MAL   Aphyosemion (Mesoaphyosemion) maculatum
MAM   Fundulopanchax (Pauciradius) marmoratus
MAN   Aplocheilus panchax marginatus
MAR   Epiplatys (=SPI) marnoi
MAT   Epiplatys (=FAS) matlocki
MAY   Nothobranchius mayeri
MAZ   Rivulus (Anablepsoides) mazaruni
MBU   Rivulus (Laimosemion) mabura
MCB   Simpsonichthys (Hypsolebias) macaubensis
MCO   Fundulus macrolepidota
MCP   Rivulus (Anablepsoides) micropus
MCR   Cynolebias microphthalmus
MCS   Aplocheilichthys macrurus
MCT   Austrolebias (=BLL) maculatus
MDI   Fundulus macdonaldi
MDT   Rivulus (Melanorivulus) modestus
MDU   Orestias mundus
MEA   Aphanius (Anatolichthys) maeandricus
MEE   Fundulus meeki
MEI   Aphyosemion meinkeni
MEJ   Rivulus (Oditichthys) mejiai
MEL   Aphyosemion melantereon
MEN   Aphanius (Paraphanius) mento
MES   Epiplatys (Epiplatys) mesogramma
MET   Aphyosemion (=CON) melanopteron
MEX   Profundulus mexicanus
MFC   Fundulus multifaciata
MGD   Rivulus (Cynodonichthys) magdalenae
MGF   Simpsonichthys (Hypsolebias) magnificus
MGR   Rivulus (Melanorivulus) megaroni
MGS   Simpsonichthys (Hypsolebias) marginatus
MIC   Aphyosemion (=CAM) microstomus
MIK   Aphyosemion (=WAC) wachtersi mikeae
MIL   Nothobranchius (Cynobranchius) microlepis
MIM   Aphyosemion (Mesoaphyosemion) mimbon
MIO   Cyprinodon nevadensis mionectes
MIR   Fundulopanchax (Paraphyosemion) mirabilis
MIT   Aphanius (Anatolichthys) meridionalis
MKU   Nothobranchius (Nothobranchius) mkuziensis
MLB   Aphyosemion (Chromaphyosemion) malumbresi
MLG   Aphyosemion (Chromaphyosemion) melanogaster
MLI   Rivulus (Cynodonichthys) milesii
MLO   Aphyosemion (Chromaphyosemion) melinoeides
MLZ   Nothobranchius (Zononothobranchius) milvertzi
MNA   Rivulus (Laimosemion) manaensis
MND   Aphyosemion (Mesoaphyosemion) mandoroense
MNE   Aphyosemion (Mesoaphyosemion) montealenense
MNG   Aphyosemion (Iconisemion) mengilai
MNK   Rivulus (Cynodonichthys) monikae
MNO   Fundulus menona
MNS   Moema manuensis
MNT   Aplocheilus melanopterus
MNU   Orestias minimus
MOA   Crenichthys baileyi moapae
MOE   Fundulopanchax (Paraphyosemion) moensis
MON   Epiplatys (Lycocyprinus) dageti monroviae
MOO   Orestias mooni
MPH   Trigonectes macrophthalmus
MPI   Cyprinodon macrolepis
MPO   Orestias multiporus
MPT   Aphanius mesopotamicus
MRG   Aphyosemion (Mesoaphyosemion) marginatum
MRI   Rivulus (Atlantirivulus) maricensis
MRM   Rivulus (Kryptolebias) marmoratus
MRT   Simpsonichthys (Hypsolebias) martinsi
MSS   Nothobranchius (Zononothobranchius) malaissei
MTC   Rivulus (Anablepsoides) monticola
MTD   Aphanius (Paraphanius) mentoides
MTM   Rivulus (Cynodonichthys) montium
MTS   Aphanius (Anatolichthys) marassantensis
MUC   Aphyosemion multicolor
MUD   Fundulus mudfish
MUF   Epiplatys (Epiplatys) multifasciatus
MUL   Orestias mulleri
MUN   Nothobranchius spec. Munyenze
MUR   Simpsonichthys (Hypsolebias) multiradiatus
MUS   Aphyosemion (Aphyosemion) musafirii
MUT   Orestias minutus
MWA   Nothobranchius spec. Mwazuma
MYE   Rivulus (Cynodonichthys) myersi
Sorted by Genus
MTD   Aphanius (Paraphanius) mentoides
MIT   Aphanius (Anatolichthys) meridionalis
MEN   Aphanius (Paraphanius) mento
MPT   Aphanius mesopotamicus
MTS   Aphanius (Anatolichthys) marassantensis
MEA   Aphanius (Anatolichthys) maeandricus

MNE   Aphyosemion (Mesoaphyosemion) montealenense
MND   Aphyosemion (Mesoaphyosemion) mandoroense
MEL   Aphyosemion melantereon
MNG   Aphyosemion (Iconisemion) mengilai
MLO   Aphyosemion (Chromaphyosemion) melinoeides
MUC   Aphyosemion multicolor
MEI   Aphyosemion meinkeni
MLG   Aphyosemion (Chromaphyosemion) melanogaster
MLB   Aphyosemion (Chromaphyosemion) malumbresi
MIM   Aphyosemion (Mesoaphyosemion) mimbon
MIK   Aphyosemion (=WAC) wachtersi mikeae
MIC   Aphyosemion (=CAM) microstomus
MAG   Aphyosemion (=CHR) margaretae
MET   Aphyosemion (=CON) melanopteron
MAL   Aphyosemion (Mesoaphyosemion) maculatum
MUS   Aphyosemion (Aphyosemion) musafirii
MRG   Aphyosemion (Mesoaphyosemion) marginatum

MCS   Aplocheilichthys macrurus

MAN   Aplocheilus panchax marginatus
MNT   Aplocheilus melanopterus
MAI   Aplocheilus panchax mattei

MAE   Archiaphyosemion (Nimbapanchax) maeseni


MAB   Austrolebias (=AFI) mabelis
MCT   Austrolebias (=BLL) maculatus



MOA   Crenichthys baileyi moapae

MCR   Cynolebias microphthalmus


MIO   Cyprinodon nevadensis mionectes
MPI   Cyprinodon macrolepis
MAA   Cyprinodon maya


MUF   Epiplatys (Epiplatys) multifasciatus
MON   Epiplatys (Lycocyprinus) dageti monroviae
MAR   Epiplatys (=SPI) marnoi
MES   Epiplatys (Epiplatys) mesogramma
MAC   Epiplatys (=SIN) macrostigma
MAT   Epiplatys (=FAS) matlocki

MAM   Fundulopanchax (Pauciradius) marmoratus
MIR   Fundulopanchax (Paraphyosemion) mirabilis
MOE   Fundulopanchax (Paraphyosemion) moensis

MUD   Fundulus mudfish
MEE   Fundulus meeki
MDI   Fundulus macdonaldi
MCO   Fundulus macrolepidota
MFC   Fundulus multifaciata
MNO   Fundulus menona






MNS   Moema manuensis

MAY   Nothobranchius mayeri
MSS   Nothobranchius (Zononothobranchius) malaissei
MLZ   Nothobranchius (Zononothobranchius) milvertzi
MKU   Nothobranchius (Nothobranchius) mkuziensis
MWA   Nothobranchius spec. Mwazuma
MIL   Nothobranchius (Cynobranchius) microlepis
MUN   Nothobranchius spec. Munyenze


MUL   Orestias mulleri
MPO   Orestias multiporus
MUT   Orestias minutus
MNU   Orestias minimus
MDU   Orestias mundus
MOO   Orestias mooni




MEX   Profundulus mexicanus


MTM   Rivulus (Cynodonichthys) montium
MYE   Rivulus (Cynodonichthys) myersi
MAZ   Rivulus (Anablepsoides) mazaruni
MTC   Rivulus (Anablepsoides) monticola
MBU   Rivulus (Laimosemion) mabura
MCP   Rivulus (Anablepsoides) micropus
MDT   Rivulus (Melanorivulus) modestus
MEJ   Rivulus (Oditichthys) mejiai
MGD   Rivulus (Cynodonichthys) magdalenae
MAH   Rivulus (Laimosemion) mahdiaensis
MGR   Rivulus (Melanorivulus) megaroni
MLI   Rivulus (Cynodonichthys) milesii
MNA   Rivulus (Laimosemion) manaensis
MNK   Rivulus (Cynodonichthys) monikae
MRM   Rivulus (Kryptolebias) marmoratus
MRI   Rivulus (Atlantirivulus) maricensis

MGS   Simpsonichthys (Hypsolebias) marginatus
MRT   Simpsonichthys (Hypsolebias) martinsi
MCB   Simpsonichthys (Hypsolebias) macaubensis
MUR   Simpsonichthys (Hypsolebias) multiradiatus
MGF   Simpsonichthys (Hypsolebias) magnificus

MPH   Trigonectes macrophthalmus

Sorted by Species
MOA   baileyi moapae, Crenichthys
MON   dageti monroviae, (Lycocyprinus) Epiplatys
MAF   gardneri mamfense, (Paraphyosemion) Fundulopanchax
MAB   mabelis, (=AFI) Austrolebias
MBU   mabura, (Laimosemion) Rivulus
MCB   macaubensis, (Hypsolebias) Simpsonichthys
MDI   macdonaldi, Fundulus
MCO   macrolepidota, Fundulus
MPI   macrolepis, Cyprinodon
MPH   macrophthalmus, Trigonectes
MAC   macrostigma, (=SIN) Epiplatys
MCS   macrurus, Aplocheilichthys
MAL   maculatum, (Mesoaphyosemion) Aphyosemion
MCT   maculatus, (=BLL) Austrolebias
MEA   maeandricus, (Anatolichthys) Aphanius
MAE   maeseni, (Nimbapanchax) Archiaphyosemion
MGD   magdalenae, (Cynodonichthys) Rivulus
MGF   magnificus, (Hypsolebias) Simpsonichthys
MAH   mahdiaensis, (Laimosemion) Rivulus
MSS   malaissei, (Zononothobranchius) Nothobranchius
MLB   malumbresi, (Chromaphyosemion) Aphyosemion
MNA   manaensis, (Laimosemion) Rivulus
MND   mandoroense, (Mesoaphyosemion) Aphyosemion
MNS   manuensis, Moema
MTS   marassantensis, (Anatolichthys) Aphanius
MAG   margaretae, (=CHR) Aphyosemion
MRG   marginatum, (Mesoaphyosemion) Aphyosemion
MGS   marginatus, (Hypsolebias) Simpsonichthys
MRI   maricensis, (Atlantirivulus) Rivulus
MRM   marmoratus, (Kryptolebias) Rivulus
MAM   marmoratus, (Pauciradius) Fundulopanchax
MAR   marnoi, (=SPI) Epiplatys
MRT   martinsi, (Hypsolebias) Simpsonichthys
MAT   matlocki, (=FAS) Epiplatys
MAA   maya, Cyprinodon
MAY   mayeri, Nothobranchius
MAZ   mazaruni, (Anablepsoides) Rivulus
MEE   meeki, Fundulus
MGR   megaroni, (Melanorivulus) Rivulus
MEI   meinkeni, Aphyosemion
MEJ   mejiai, (Oditichthys) Rivulus
MLG   melanogaster, (Chromaphyosemion) Aphyosemion
MET   melanopteron, (=CON) Aphyosemion
MNT   melanopterus, Aplocheilus
MEL   melantereon, Aphyosemion
MLO   melinoeides, (Chromaphyosemion) Aphyosemion
MNG   mengilai, (Iconisemion) Aphyosemion
MNO   menona, Fundulus
MEN   mento, (Paraphanius) Aphanius
MTD   mentoides, (Paraphanius) Aphanius
MIT   meridionalis, (Anatolichthys) Aphanius
MES   mesogramma, (Epiplatys) Epiplatys
MPT   mesopotamicus, Aphanius
MEX   mexicanus, Profundulus
MIL   microlepis, (Cynobranchius) Nothobranchius
MCR   microphthalmus, Cynolebias
MCP   micropus, (Anablepsoides) Rivulus
MIC   microstomus, (=CAM) Aphyosemion
MLI   milesii, (Cynodonichthys) Rivulus
MLZ   milvertzi, (Zononothobranchius) Nothobranchius
MIM   mimbon, (Mesoaphyosemion) Aphyosemion
MNU   minimus, Orestias
MUT   minutus, Orestias
MIR   mirabilis, (Paraphyosemion) Fundulopanchax
MKU   mkuziensis, (Nothobranchius) Nothobranchius
MDT   modestus, (Melanorivulus) Rivulus
MOE   moensis, (Paraphyosemion) Fundulopanchax
MNK   monikae, (Cynodonichthys) Rivulus
MNE   montealenense, (Mesoaphyosemion) Aphyosemion
MTC   monticola, (Anablepsoides) Rivulus
MTM   montium, (Cynodonichthys) Rivulus
MOO   mooni, Orestias
MUD   mudfish, Fundulus
MUL   mulleri, Orestias
MUC   multicolor, Aphyosemion
MFC   multifaciata, Fundulus
MUF   multifasciatus, (Epiplatys) Epiplatys
MPO   multiporus, Orestias
MUR   multiradiatus, (Hypsolebias) Simpsonichthys
MDU   mundus, Orestias
MUN   Munyenze, spec. Nothobranchius
MUS   musafirii, (Aphyosemion) Aphyosemion
MWA   Mwazuma, spec. Nothobranchius
MYE   myersi, (Cynodonichthys) Rivulus
MIO   nevadensis mionectes, Cyprinodon
MAN   panchax marginatus, Aplocheilus
MAI   panchax mattei, Aplocheilus
MIK   wachtersi mikeae, (=WAC) Aphyosemion
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Richard J. Sexton